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Vorticity inversion at higher resolution (note, this does not require space-filling like the buoyancy field does).
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Enhanced (stochastic) merging and splitting
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Parcels with a dilution factor, which slowly grow in volume, keeping product volume*dilution. Merging based on dilution as well as other factors. Dilution/expansion rate informed by physics of mixing.
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Parcel nudging to gridded field values with time-scale representing subgrid mixing.
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Andy Visser's book on zooplankton ecology (! Chapter 4), provides a nice perspective to mixing, I think.
https://www.researchgate.net/publication/280883339_Small_wet_rational_Individual_based_zooplankton_ecology -
See also figure 5 in http://authors.library.caltech.edu/1349/1/DIMjfm00.pdf
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